The paragraph is talking about Universities as strange institutions. 'R' will be the first sentence as the words 'historical roots' highlight its origin. PS will come together and P will be the next sentence as 'traditionally' points towards the period after its inception. 'S' comes after P showing the medieval period of universities. The correct sequence will be 'RPSQ'. Naturally, 'Q' will be the last sentence of the passage. So, option (d) is correct.